Abstract

Cutaneous injury triggers sensory neurons that elicit pain or itch and modulate tissue repair. A subset of these neurons expresses the Mas-related G protein-coupled receptor MrgprD, which is selectively activated by β alanine. However, the physiological source of this endogenous metabolite has remained elusive for two decades. Here, we unexpectedly identify apoptotic cells as a physiologically relevant endogenous source of β alanine. Through an unbiased screen of G protein-coupled receptors (GPCRs) activated by metabolites released from dying cells, we detected robust activation of MrgprD. Mass spectrometry and functional assays confirmed that early-stage apoptotic cells release β alanine via the caspase-mediated opening of Pannexin 1 channels. Because MrgprD-expressing neurons innervate the epidermis, we investigated the skin and identified apoptotic keratinocytes as a potential local source of β alanine. Using three distinct models of skin perturbation, we demonstrate that MrgprD signaling directly modulates cutaneous responses. Collectively, these findings uncover a previously unrecognized signaling axis linking apoptotic metabolites to sensory neuron activation, local tissue injury responses, and the regulation of inflammation.

Committee Chair

Kodi Ravichandran

Committee Members

Claudia Han; Kendall Blumer; Marco Colona; Steven VanDyken; Xiaoxiao Wan

Degree

Doctor of Philosophy (PhD)

Author's Department

Biology & Biomedical Sciences (Molecular Cell Biology)

Author's School

Graduate School of Arts and Sciences

Document Type

Dissertation

Date of Award

8-13-2026

Language

English (en)

Available for download on Thursday, August 12, 2027

Included in

Biology Commons

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