ORCID

https://orcid.org/0000-0001-6511-5480

Date of Award

12-20-2023

Author's School

Graduate School of Arts and Sciences

Author's Department

Biology & Biomedical Sciences (Biochemistry)

Degree Name

Doctor of Philosophy (PhD)

Degree Type

Dissertation

Abstract

Intrinsically disordered regions (IDRs) are protein domains or regions that lack a fixed three- dimensional structure. Within IDRs, regions and residues can engage in intramolecular interactions with other parts of the same polypeptide or intermolecular interactions with other biomolecules. This dissertation focuses on how the “context” of an IDR can influence the interactions it can undergo. Specifically, how the identity, chemistry, and position of interacting residues in IDRs tune IDR conformational behavior, the propensity of IDRs to self-assemble via phase separation, and how these three properties are tuned by the solution environment. These concepts are explored through a combination of bioinformatic sequence analysis, all-atom (CAMPARI) and coarse grain (PIMMS + LAMMPS) simulations, and through integral wet lab collaborations. Following an initial introduction of IDRs and the possible interactions they can undergo, explored are four key themes: Chapter 3-6 explore new bioinformatic methods for analyzing and characterizing IDRs enabling a more accessible and contextualized understanding of IDR conformational properties. Subsequently, the idea of sequence “context” is then explored through a set of collaborations showing how the distributions and clustering of amino acids can affect IDR conformation and condensate properties. The chemical solution “context” of an IDR is then introduced via a method for quantifying the sensitivity of IDR, and is explored through a discussion on how the solution environment of an IDR affects its conformational behavior and capacity undergo phase separation. Finally, Chapters 9 & 10 present mechanisms for identifying amino acid sequence features that delineate the presence of interacting residues in an IDR sequence and then also explores the idea of “context” relative to how IDR interactions with other proteins and RNA can promote phase separation.

Language

English (en)

Chair and Committee

Alex Holehouse

Available for download on Sunday, December 20, 2026

Included in

Biophysics Commons

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